In a world of six billion other faces, your face is unique. It reveals your personality, your genetic and cultural identity and it can be read like a book. Family resemblances, facial recognition and the purpose of the face and its features are all part of a story that begins in the oceans five hundred million years ago.We conclude with the multi-racial face of the future and the pioneering work of surgeons in Kentucky who are preparing for the world’s first facial transplant.